The Exchange mailboxes task when run against Exchange 2010 servers has the following warning for numerous mailboxes: "The mailbox could not be associated to a store in the database.
When viewing the list of tasks that ran for the Default Gathering you can see that both the Exchange Information Stores and Exchange Databases tasks ran against the Exchange 2010 servers
In addition, the reports may have only partial data for the mailboxes reported in the warning.
Gathering corruption. The Exchange Information Stores task is not supposed to run against Exchange 2010 servers, only the Exchange Databases task.
Delete the Default Gathering and recreate it.